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In 1937, a Chinese martial artist returns to Shanghai to find his teacher dead and his school harassed by the Japanese

Gameras ties to humanity have been severed as the Gyaos re emerge Nagamine and Asagi investigate as an orphaned girl named Ayana discovers a new creature Named Iris, Ayana seeks revenge on Gamera for the death of her family

An ornithologist investigates reports of a monstrous new species of bird just as a teenage girl is gifted an amulet found on mysterious atoll As the creatures begin to attack, an ancient guardian with a bond to the girl emerges